Like arthropods, mollusks gained a foothold during the Cambrian period and eventually branched out into a profusion of divergent forms, including (clockwise from left) a modern lettuce sea slug, chiton, tusk shell, keyhole limpet, and spotted sea hare. Nectocaris pteryx, the 508-million-year-old fossil preserved in the Burgess Shale at right, sports several features seen today in squid, octopuses, and other creatures in the cephalopod class of mollusks. The shared traits include tentacles, eyes, and below them, a funnel used for propulsion. Most early mollusks lived on the ocean bottom, but Nectocaris had specialized to wander throughout the water column.…
